Jonathan joined the lab in October 2009. He completed his Ph.D. in Genetics at Stanford University in 2009 and his B.S. in Biology at Humboldt State University in 2004. He is currently an NIH postdoctoral fellow supported by the NIGMS. His research is about what makes centromeres distinct from other parts of chromosomes and how they function as stable sites of kinetochore assembly. In particular, he is studying centromere encoded RNA, centromere chromatin structure, and dynamics of centromere positioning. In addition to centromeres, he is also studying interactions between intergenic and genic chromatin, especially in regard to RNA-based chromatin regulation.